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Résumé :
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In 1994 a survey of calling Tengmalm's Owls was conducted in a 28 km2 forest area around Hanstedt Hills in the north-eastern part of the Nature Reserve "Luneburger Heide" (Lower Saxony, Germany). A high density of 1,3 calling males per km2 was found. This is due to several reasons: (1) The forests contain several areas with trees more than 80 years old (beech, spruce, pine). In these old trees the Black Woodpecker has constructed many holes, which can be used by Tengmalm's Owl for nesting. (2) The strong storms of the years 1990-1992 have generated several large windfall areas, which are optimal foraging areas for Tengmalm's Owl. (3) The Tawny Owl, an important predator of Tengmalm's Owls, lives predominantly in the surroundings of small villages, not inside the contiguous forest, because the winter climate is rather hard in the Hanstedt Hills and the occurrence of small mammals is, probably, limited on the poorer soils of the hills. In the long term, the spread of Tengmalm's Owl in the northern German lowlands was a consequence of reforestation and the concurrent spread of Black Woodpecker, on whose cavities the owl depends.
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