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Résumé :
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A comparative study of the habitat use of Nuthatch, Great and Middle Spotted Woodpeckers was carried out in two managed, 100-165 years old beech forests of 40 ha each in the Schorfheide-Chorin Biosphere Reserve in the north-east of Brandenburg. As reference areas, the strict nature reserves "Heilige Hallen" (25 ha) in the south of Mecklenburg-Vorpommern and "Fauler Ort" (14 ha) in the northern part of the Biosphere Reserve were chosen. Both are up to 350-year-old beech forests which have not been managed for more than 150 years. Data were gathered from March to July and from November to December 2000. Foraging behaviour of the three species and the use of structures and substrates were studied. Of special interest was the question of the varying use of resources in nearly pure beech forests by the Middle Spotted Woodpecker, which was formerly thought to be mainly dependent on oaks. When searching for animal food, both woodpecker species were disproportionally using dead wood of beech trees, P. major seasonally used mostly fallen, P. medius standing dead wood. Beside the tree-stumps, the almost dead beeches play a crucial role in the near-natural beech forests. In the managed forests, the small proportion of oaks was used predominantly, P. medius even used the oaks seasonally by a majority. When searching for food on living substrate in the unmanaged (near-natural) forests, P. medius used exclusively bark patches with a rough, rich-structured surface ("bark patches of malfunction"). Such structures like fallen dead wood, dead and almost dead beeches preferred by P. major and Sitta europaea obviously enable these species to inhabit the natural beech forests with clearly higher abundances. In case of P. medius, the high proportion of standing dead wood (also as a part of living trees) and the richly structured surface of mature beech trees allow the occurrence in pure beech forests. It can be assumed, therefore that in former times lowland beech forests were widely inhabited by the Middle Spotted Woodpecker. Specific silvicultural recommendations for the support of P. medius in managed beech forests are drawn.
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